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Öğe The alien ascidian Styela clava now invading the Sea of Marmara (Tunicata: Ascidiacea)(Pensoft Publ, 2016) Cinar, Melih ErtanDuring the implementation of a large project aimed to investigate the benthic community structures of the Sea of Marmara, specimens of the invasive ascidian species Styela clava were collected on natural substrata (rocks) at 10 m depth at one locality (Karamursel) in Izmit Bay. The specimens were mature, containing gametes, indicating that the species had become established in the area. The Sea of Marmara seems to provide suitable conditions for this species to survive and form proliferating populations.Öğe ALien Biotic IndEX (ALEX) - A new index for assessing impacts of alien species on benthic communities(Pergamon-Elsevier Science Ltd, 2014) Cinar, Melih Ertan; Bakir, KeremBiotic indices are mainly aimed at assessing levels of deterioration caused by chemical or organic pollution. However, no biotic index to date has been developed to detect impacts of alien species on benthic communities. In this paper, a new biotic index, namely ALEX, is proposed to address the objectives of the Water Framework Directive and was tested in Mersin Bay (Levantine Sea, Turkey). Species were divided into four biogeographic groups, namely native species, casual species, established species and invasive species, and the metric considers the relative importance of these groups in samples. The index classified the ecological status of some stations which are shallow, and close to harbor and river mouths as bad or poor in February and October. The ALEX values were positively and significantly correlated with total nitrogen, silicate and silt percentage in sediment, and negatively correlated with depth and the distance from the harbor. (C) 2014 Elsevier Ltd. All rights reserved.Öğe Alien polychaete species (Annelida: Polychaeta) on the southern coast of Turkey (Levantine Sea, eastern Mediterranean), with 13 new records for the Mediterranean Sea(Taylor & Francis Ltd, 2009) Cinar, Melih ErtanThe examination of benthic material collected from different depths and habitats along the southern coast of Turkey (Levantine Sea, eastern Mediterranean) in September and October 2005 revealed 30 alien polychaete species belonging to 15 families. Thirteen species (Pisione guanche, Linopherus canariensis, Onuphis eremita oculata, Lumbrineris perkinsi, Dorvillea similis, Timarete caribous, Pherusa parmata, Pherusa saldanha, Streblosoma comatus, Polycirrus twisti, Laonome triangularis, Branchiomma bairdi and Janua steueri) are new to the Mediterranean fauna, 24 species are new to the Turkish fauna and 27 species are new to the Levantine coast of Turkey. The specimens collected from the Levantine coast of Turkey and northern Cyprus that were previously identified as Linopherus acarunculata and Branchiomma boholense are reidentified as L. canariensis and B. bairdi, respectively. In the light of present knowledge, the previous reports of Lumbrineris inflata and Streblosoma hesslei from the Mediterranean (Italian coast) could refer to the species L. perkinsi and S. comatus, respectively. The species new to the Mediterranean Sea are redescribed and discussed.Öğe Alien polychaete species worldwide: current status and their impacts(Cambridge Univ Press, 2013) Cinar, Melih ErtanThis paper reviews the alien polychaete species in the world's oceans and their impacts on the marine ecosystems and humans. A total of 292 polychaete species belonging to 164 genera and 39 families have been transported around the world's oceans with human-mediated assistance. Almost 15% of total number of known polychaete genera and 3.4% of total number of species are included in this phenomenon. A total of 180 species have become established in the world's oceans and 31 species (casual species) have a potential to establish viable populations in a new location. The most speciose genera are Hydroides (16 species) and Polydora (16 species), both accounting for 10% of the total number of alien species. The families Spionidae (53 species) and Serpulidae (46 species) have the highest number of alien species. The Mediterranean Sea (134 species), and the coasts of the Hawaii Islands (47 species) and the USA Pacific (34 species) have been intensively invaded by alien polychaetes. The origins of alien species vary among regions. Alien polychaete species in the Mediterranean Sea mostly originated from the Red Sea and Indo-Pacific areas. Benthic habitats of the areas between 40 degrees N and 40 degrees S were colonized by polychaetes mostly originating from other tropical and subtropical regions. The Suez Canal and shipping are the major vectors for species introductions. Some species imported and exported as fishing baits have become established at non-native localities. The invasive polychaete species have greatly altered habitat structures in some areas, restructured the food webs, and created important economic problems.Öğe Alien Species Threat across Marine Protected Areas of Turkey-An Updated Inventory(Mdpi, 2021) Bilecenoglu, Murat; Cinar, Melih ErtanThis study presents the first comprehensive assessment of alien species occurrences within the selected 11 Marine Protected Areas (MPAs) located on the Aegean and Levantine coasts of Turkey. The inventory includes a total of 289 species belonging to 15 phyla, in which lowest and highest diversities were observed in Saros Bay MPA (27 species, northern Aegean Sea) and Fethiye-Gocek Bay MPA (150 species, northwest Levantine Sea), respectively. Alien species distributions that were revealed in protected areas located in the southern Aegean and Levantine Seas were 56.9% similar (based on presence vs. absence data), while northern Aegean sites formed another distinct group. According to the breakdown of major phyla through the entire study areas, Mollusca had the highest alien diversity (22.1% of alien species), followed by Actinopterygii (19.0%), Arthropoda (15.2%) and Annelida (13.5%). Casual aliens were represented by very low proportions in each MPA, proving that most species were already established in the region, with a significant proportion of invasive species. Regardless of the localities, the majority of the species originated from the Red Sea, whose primary pathway of introduction is the corridor, the Suez Canal. In the absence of effective management actions against bioinvasions, MPAs located along the Turkish coastline do not currently seem to provide any protection, revealing a large conservation gap to be filled.Öğe Annelids (Polychaeta and Oligochaeta) from the Sea of Marmara, with descriptions of five new species(Taylor & Francis Ltd, 2011) Cinar, Melih Ertan; Dagli, Ertan; Acik, SerminThis paper deals with annelids (Oligochaeta and Polychaeta) collected in the Sea of Marmara between 2006 and 2010 at depths from 0 to 66 m. A total of five oligochaete and 198 polychaete species were found. Five polychaete species, namely Prosphaerosyllis marmarae sp. nov., Levinsenia demiri sp. nov., Levinsenia kosswigi sp. nov., Levinsenia marmarensis sp. nov. and Levinsenia tribranchiata sp. nov. are new to science, and five oligochaete and 84 polychaete species are new to the fauna of the Sea of Marmara. A list of species found in the region and their maximum densities are presented at depth intervals. The present material includes six alien polychaete species, Paraprionospio coora, Polydora cornuta, Prionospio (Minuspio) pulchra, Pseudopolydora paucibranchiata, Chaetozone corona and Metasychis gotoi, of which the latter four species were new records for the region.Öğe Bioeroding (boring) polychaete species (Annelida: Polychaeta) from the Aegean Sea (eastern Mediterranean)(Cambridge Univ Press, 2021) Cinar, Melih Ertan; Dagli, ErtanThe present study reports polychaetes that bore into limestone rocks along the east coast of the Aegean Sea (eastern Mediterranean). Rock materials were collected at two depth intervals (0-5 and 5-10 m) at 15 stations in four localities of Ildiri Bay. A total of 276 specimens belonging to 12 species and four families (Eunicidae, Spionidae, Cirratulidae and Sabellidae) were recorded. Specimens belonging to Dodecaceria and Pseudopotamilla were identified at the genus level, because they differ from described species, were few in number or were in poor condition. Dipolydora giardia is a new species to the marine fauna of Turkey. The most dominant and frequent family in the area was Eunicidae, followed by Spionidae. Lysidice ninetta and L. margaritacea comprised 59% of the total number of individuals. The number of species and individuals, and the diversity index did not change with regard to depth or locality. Two species assemblages were found in the area, mainly formed by Dipolydora and Lysidice species. The Lessepsian species, Palola valida, which is a new record for the Aegean Sea, occurred abundantly at the study sites, posing a risk of damage to limestone rocks in the Mediterranean Sea. The morphological features of the species identified at the generic level and the burrow structure of these species are presented. The burrow shapes of Palola siciliensis and P. valida were described for the first time in the present study; they constructed complicated galleries, including more than four entrances.Öğe Brittle stars (Echinodermata: Ophiuroidea) from the southern coast of Turkey (eastern Mediterranean): new records and revision of Amphiodia obtecta Mortensen, 1940(Magnolia Press, 2010) Stohr, Sabine; Cinar, Melih Ertan; Dagli, ErtanBrittle stars were collected on the southern coast of Turkey in 2005 and 2009. The most abundant species found (up to 420 ind. m(-2)) was Amphiodia obtecta. The taxonomic status of this species was unclear and is revised here. This is the first record of the genus Amphiodia in the Mediterranean Sea; it probably originates from the Red Sea, since it is absent from the entire Atlantic Ocean. Among the other species found, Ophiactis macrolepidota (previously reported as O. parva in the Mediterranean) is a new record for the marine fauna of Turkey.Öğe Checklist of Annelida from the coasts of Turkey(Tubitak Scientific & Technical Research Council Turkey, 2014) Cinar, Melih Ertan; Dagli, Ertan; Kurt Sahin, GuleyThe compilation of papers on marine annelids along the coasts of Turkey together with new records of species (24 species) presented in this study yielded a total of 721 species belonging to 2 classes (Polychaeta and Clitellata), 60 families, and 352 genera. Polychaeta were represented by 705 species, Oligochaeta by 13 species, and Hirudinea by 3 species. Syllidae (119 species) and Serpulidae (56 species) were the species-rich polychaete families. The majority of annelid species were benthic (691 species), 14 species were pelagic, and 3 species (leeches) were parasitic. Thirteen polychaete species were excluded from the species inventory. The Aegean Sea had the highest number of species (559 species), followed by the Levantine Sea (459 species) and the Sea of Marmara (398 species). The hot spot areas for the species diversity were Izmir Bay, Mersin Bay, the southwest part of the Sea of Marmara, and Sinop Peninsula, where intense scientific efforts have been carried out. A total of 75 alien polychaete species were reported from the regions, 22 of which were classified as invasive species. The annelid species were generally encountered in soft substrata of the shallow-water benthic environments, whereas only 9 species were reported from depths deeper than 600 m.Öğe Checklist of Cnidaria and Ctenophora from the coasts of Turkey(Tubitak Scientific & Technical Research Council Turkey, 2014) Cinar, Melih Ertan; Yokes, Mehmet Baki; Acik, Sermin; Bakir, Ahmet KeremThis paper presents the actual status of species diversity of the phyla Cnidaria and Ctenophora along the Turkish coasts of the Black Sea, the Sea of Marmara, the Aegean Sea, and the Levantine Sea. A total of 195 cnidarian species belonging to 5 classes (Hydrozoa, Cubozoa, Scyphozoa, Staurozoa, and Anthozoa) have been determined in these regions. Eight anthozoan species (Arachnanthus oligopodus, Bunodactis rubripunctata, Bunodeopsis strumosa, Corynactis viridis, Halcampoides purpureus, Sagartiogeton lacerates, Sagartiogeton undatus, and Pachycerianthus multiplicatus) are reported for the first time as elements of the Turkish marine fauna in the present study. The highest number of cnidarian species (121 species) was reported from the Aegean Sea, while the lowest (17 species) was reported from the Black Sea. The hot spot areas for cnidarian diversity are the Prince Islands, Istanbul Strait, Izmir Bay, and Datca Peninsula, where relatively intensive scientific efforts have been carried out. Regarding ctenophores, 7 species are distributed along the Turkish coasts, 5 of which were reported from the Black Sea. A total of 16 alien cnidarian and 2 ctenophore species were determined in the regions. Two species (Sagartiogeton laceratus and Pachycerianthus multiplicatus) are new alien species for the Mediterranean Sea and could have been introduced to the northern part of the Sea of Marmara and Iskenderun Bay, areas from which these species are recorded, by ships from the North-East Atlantic.Öğe Checklist of the phyla Platyhelminthes, Xenacoelomorpha, Nematoda, Acanthocephala, Myxozoa, Tardigrada, Cephalorhyncha, Nemertea, Echiura, Brachiopoda, Phoronida, Chaetognatha, and Chordata (Tunicata, Cephalochordata, and Hemichordata) from the coasts of Turkey(Tubitak Scientific & Technical Research Council Turkey, 2014) Cinar, Melih ErtanIn this paper, the current status of the species diversity of 13 phyla, namely Platyhelminthes, Xenacoelomorpha, Nematoda, Acanthocephala, Myxozoa, Tardigrada, Cephalorhyncha, Nemertea, Echiura, Brachiopoda, Phoronida, Chaetognatha, and Chordata (invertebrates, only Tunicata, Cephalochordata, and Hemichordata) along the coasts of Turkey is reviewed. Platyhelminthes was represented by 186 species, Chordata by 64 species, Nemertea by 26 species, Nematoda by 20 species, Xenacoelomorpha by 11 species, Chaetognatha by 10 species, Acanthocephala by 9 species, Brachiopoda and Phoronida by 4 species, Myxozoa and Tradigrada by 2 species, and Cephalorhyncha and Echiura by 1 species. Two platyhelminth (Planocera cf. graffi and Prostheceraeus vittatus), 2 nemertean (Drepanogigas albolineatus and Tubulanus superbus), 1 phoronid (Phoronis australis), and 2 ascidian (Polyclinella azemai and Ciona roulei) species are being newly reported for the first time from the coasts of Turkey. Four tunicate (Symplegma brakenhielmi, Microcosmus exasperatus, Herdmania momus, and Phallusia nigra) and 1 chaetognath (Ferosagitta galerita) species were classified as alien species in the region.Öğe CIGESMED for divers: Establishing a citizen science initiative for the mapping and monitoring of coralligenous assemblages in the Mediterranean Sea(Pensoft Publ, 2016) Gerovasileiou, Vasilis; Dailianis, Thanos; Panteri, Emmanouela; Michalakis, Nikitas; Gatti, Giulia; Sini, Maria; Dimitriadis, Charalampos; Issaris, Yiannis; Salomidi, Maria; Filiopoulou, Irene; Dogan, Alper; d'Avray, Laure Thierry de Ville; David, Romain; Cinar, Melih Ertan; Koutsoubas, Drosos; Feral, Jean-Pierre; Arvanitidis, ChristosBackground Over the last decade, inventorying and monitoring of marine biodiversity has significantly benefited from the active engagement of volunteers. Although several Citizen Science projects concern tropical reef ecosystems worldwide, none of the existing initiatives has yet specifically focused on their Mediterranean equivalents. Mediterranean coralline reefs, known as "coralligenous", are bioherms primarily built by calcifying rhodophytes on hard substrates under dim-light conditions; they are considered hotspots of biodiversity and are extremely popular among divers due to their complex structure, conspicuous biological wealth and high aesthetic value. Nevertheless, data on their distribution, structure and conservation status is lacking for several Mediterranean areas while they are vulnerable to an increasing number of threats. New information In the framework of CIGESMED SeasEra (ERAnet) project a specialized Citizen Science project was launched, aiming to engage enthusiast divers in the study and monitoring of Mediterranean coralligenous assemblages through the gathering of basic information regarding their spatial occurrence, assemblage structure and associated pressures or threats. For its active implementation, a data collection protocol and a multilingual website were developed, comprising an educational module and a data submission platform. Georeferenced data reporting focuses on: (a) basic topographic and abiotic features for the preliminary description of each site, and the creation of data series for sites receiving multiple visits; (b) presence and relative abundance of typical conspicuous species, as well as (c) existence of pressures and imminent threats, for the characterization and assessment of coralligenous assemblages. A variety of tools is provided to facilitate end users, while divers have the choice to report additional information and are encouraged to upload their photographs. The long-term goal is the development of an active community of amateur observers providing widespread and ecologically significant data on coralligenous assemblages.Öğe Cirrophorus species (Annelida: Polychaeta: Paraonidae) in the Sea of Marmara, with description of a new species(Magnolia Press, 2017) Erdogan-Dereli, Deniz; Cinar, Melih Ertan; Dagli, ErtanThis paper deals with specimens of Cirrophorus collected from the depths ranging from 3 to 500 m in 2012 and 2013 in the Sea of Marmara, Turkey. Among the material, one new species and two already known species were found. The new species is mainly characterized by having long and clavate notopodial postchaetal lobes in the branchial region; red speckles near nuchal organs; and a very long branch of lyrate chaetae (4 times longer than the shorter one). A recently described species, Cirrophorus nikebianchii, was found at the majority of soft-bottom stations. All species found in the area were described and discussed.Öğe Coralligenous assemblages along their geographical distribution: Testing of concepts and implications for management(Wiley, 2020) Cinar, Melih Ertan; Feral, Jean-Pierre; Arvanitidis, Christos; David, Romain; Taskin, Ergun; Sini, Maria; Katagan, TuncerThe coralligenous habitat was studied at the large Mediterranean scale, by applying a standardized, non-destructive photo-sampling protocol, developed in the framework of the CIGESMED project. the results provided evidence to support the following statements: (a) the assemblage pattern is not homogeneously distributed across the four Mediterranean ecoregions studied (biotic gradients hypothesis); and (b) the assemblage pattern does not change significantly when the information is aggregated to higher taxonomic levels (taxonomic sufficiency hypothesis). Surrogate taxonomic categories higher than species, such as genus and family, can be used to reveal the multivariate pattern of the coralligenous assemblages. Although preliminary at the pan-Mediterranean scale, these outcomes set the scene for future comparisons as more data sets become available but also for comparisons between taxonomic and functional patterns. 1.2.3.4.Öğe Current status (as of end of 2020) of marine alien species in Turkey(Public Library Science, 2021) Cinar, Melih Ertan; Bilecenoglu, Murat; Yokes, M. Baki; Ozturk, Bilal; Taskin, Ergun; Bakir, Kerem; Dogan, AlperThe 2020's update of marine alien species list from Turkey yielded a total of 539 species belonging to 18 taxonomic groups, 404 of which have become established in the region and 135 species are casual. A total of 185 new alien species have been added to the list since the previous update of 2011. The present compilation includes reports of an ascidian species (Rhodosoma turcicum) new to the marine fauna of Turkey and range extensions of six species. Among the established species, 105 species have invasive characters at least in one zoogeographic region, comprising 19% of all alien species. Mollusca ranked first in terms of the number of species (123 species), followed by Foraminifera (91 species), Pisces (80 species) and Arthropoda (79 species). The number of alien species found in seas surrounding Turkey ranged from 28 (Black Sea) to 413 (Levantine Sea). The vectoral importance of the Suez Canal diminishes when moving from south to north, accounting for 72% of species introductions in the Levantine Sea vs. only 11% of species introductions in the Black Sea. Most alien species on the coasts of Turkey were originated from the Red Sea (58%), due to the proximity of the country to the Suez Canal. Shipping activities transported 39% of alien species, mainly from the Indo-Pacific area (20%) and the Atlantic Ocean (10%). Misidentified species (such as Pterois volitans, Trachurus declivis, etc.) and species those classified as questionable or cryptogenic were omitted from the list based on new data gathered in the last decade and expert judgements. The documented impacts of invasive species on socio-economy, biodiversity and human health in the last decade as well as the legislation and management backgrounds against alien species in Turkey are presented.Öğe Description of a new fireworm, Eurythoe turcica sp nov (Polychaeta : Amphinomidae), from the Levantine coast of Turkey (eastern Mediterranean), with re-descriptions of Eurythoe parvecarunculata Horst and Amphinome djiboutiensis Gravier based on type material(Taylor & Francis Ltd, 2008) Cinar, Melih ErtanThe present study describes a new fireworm belonging to the genus Eurythoe, E. turcica sp. nov., collected from Iskenderun Bay and its vicinity (Levantine Sea, Turkey), and re-describes the closely related species E. parvecarunculata Horst, 1912 and Amphinome djiboutiensis Gravier, 1902, based on the type material. The new species is mainly characterized by having a rounded and short caruncle occupying only chaetiger 1, and dendritically branched branchiae from chaetiger 3 to the end of body. Morphological features of E. turcica are closely similar to those of E. parvecarunculata but the former species lacks the serrated capillary chaeta with a long spur on the notopodia of the median parapodia. The present study also shows that A. djiboutiensis and E. parvecarunculata, which were previously considered synonymous, are distinct species.Öğe Descriptions of two new genera, six new species and three new records of Anthozoa (Cnidaria) from the Sea of Marmara(Taylor & Francis Ltd, 2018) Ocana, Oscar; Cinar, Melih ErtanThe present study describes two new genera, six new species and three new records from the Sea of Marmara. The new genus Marmara gen. nov. belonging to the family Sagartiidae is mainly characterized by having conspicuous muscular belts along the column and the absence of suckers and tenaculi on the body wall. The genus Charisactis gen. nov. belonging to the family Condylanthidae is mainly characterized by having an atypical mesentery arrangement, single siphonoglyph and homotrichs in tentacles. The other species newly described from the Sea of Marmara belong to the families Diadumenidae (one species from deep waters), Halcampoididae (one species from shallow waters), Edwardsiidae (one species from shallow waters) and Epizoanthidae (one species from deep waters). Three species, namely Epizoanthus arenaceus (family Epizoanthidae), Rolandia coralloides (family Clavulariidae) and Virgularia sp. (family Virgulariidae) are reported for the first time from the Sea of Marmara. The external and internal anatomical features as well as cnidom structures of the species are presented and discussed with the closely related species. http://www.zoobank.org/urn:lsid:zoobank.org:pub:51DA9869-D786-4A8A-8C1F-62DC1BB67923Öğe Distribution of Eunicidae (Annelida: Polychaeta) along the Levantine coast of Turkey, with special emphasis on alien species(Springer Heidelberg, 2017) Kurt-Sahin, Guley; Cinar, Melih ErtanDistributional features of Eunicidae (Annelida, Polychaeta) along the Levantine coast of Turkey were assessed based on data derived from qualitative and quantitative samples collected in various substrata, habitat and depths (0-100 m). The examined material contained 12 species and 4644 individuals belonging to five genera (Eunice, Leodice, Lysidice, Marphysa and Palola). Marphysa was represented by the highest number of species and Lysidice by the highest number of individuals in the area. The dominant species were Lysidice ninetta, L. margaritacea and Leodice antennata; the most frequent species were Leodice antennata, Lysidice ninetta and L. unicornis among the stations. The Lessepsian migrants, Leodice antennata and Palola valida, seem to have been well established and formed dense populations (max. 725 and 100 ind.m(-2), respectively) in the area. The distribution of L. antennata seems to be related to the total inorganic nitrogen and silica concentrations in the water column. The ratio between alien and native species' abundances increased from the western to the eastern parts of the region.Öğe Distribution of polychaete species (Annelida: Polychaeta) on the polluted soft substrate of the Golden Horn Estuary (Sea of Marmara), with special emphasis on alien species(Cahiers De Biologie Marine, 2009) Cinar, Melih Ertan; Balkis, Huesamettin; Albayrak, Serhat; Dagli, Ertan; Karhan, Selahattin UensalThe present study deals with the polychaete assemblages inhabiting Polluted soft substrate of the Golden Horn Estuary (Sea of Marmara) and the importance of alien species in polychaete communities. A total of 15 species belonging to 7 families were determined among the benthic materials collected at 4 stations between the depths 4 and 40 in in December 2005. The materials were composed of species tolerant to pollution. Higher diversity and evenness index values were encountered at stations where dissolved oxygen values were relatively high and total organic carbon values were low. The material included four alien species: Desdemona ornata, Ficopomatus enigmaticus, Polydora cornuta and Streblospio gynobranchiata. The most dominant alien species in the area was P. cornuta, with 4340 individuals per square meter. Alien species comprised 46% of total polychaete populations and their distributions were strongly affected by the abiotic factors such as dissolved oxygen, total organic carbon and depth.Öğe The diversity of polychaetes (Annelida: Polychaeta) in a long-term pollution monitoring study from the Levantine coast of Turkey (Eastern Mediterranean), with the descriptions of four species new to science and two species new to the Mediterranean fauna(Taylor & Francis Ltd, 2022) Cinar, Melih Ertan; Dagli, Ertan; Erdogan-Dereli, DenizDuring a relatively long-term pollution monitoring study between 2015 and 2021 (annually between 2015 and 2019; triennially after 2019), a total of 257 polychaete species belonging to 40 families were identified at 17 soft bottom stations (depth range: 11-89 m) along the Levantine coast of Turkey. Among them, four species belonging to Hesionidae (Oxydromus digitifera sp. nov.), Orbiniidae (Leitoscoloplos mediterranea sp. nov.), Paraonidae (Paraonis lobulata sp. nov.) and Capitellidae (Barantolla cryptogenica sp. nov.) are new to science, two species belonging to Cirratulidae (Chaetozone elakata) and Ampharetidae (Anobothrus amoureuxi) are new additions to the Mediterranean fauna and 28 species are new additions to the marine fauna of Turkey. Among the families, Spionidae (29 species), Syllidae (22 species) and Paraonidae (22 species) accounted for 28% of the total number of species, and Capitellidae, Lumbrineridae and Spionidae comprised 45% of the total number of individuals. The most dominant species in the region were Lumbrineris geldiayi, Notomastus mossambicus and Prionospio saccifera. A total of 20 alien species belonging to 12 families were found in the region, with N. mossambicus and P. saccifera accounting for more than 85% of the total number of individuals of alien species. The new species are described and compared to the closely related species.
